By Scott Solomon, Rice University
The sun temple built by pharaoh Nyuserre in the Fifth Dynasty of ancient Egypt 4,500 years ago has the earliest known depiction of beekeeping. But remnants of beeswax in pottery fragments from 9,000 years ago in Turkey suggest that beekeeping has an even longer history going back to the very start of agriculture.

History of Beekeeping
It’s not hard to imagine why ancient people would want to keep bees around. Honey was one of the only available sweeteners in the ancient Near East. Beeswax also had medicinal and practical uses, for example as parts of tools and as a waterproofing agent.
Also, as modern beekeepers know, bees are relatively easy to work with. Unlike many other stinging insects, the European honeybee, Apis mellifera, is relatively reluctant to sting. That’s especially true when a bee fills up with honey, which honeybees do instinctively when they smell smoke.
While some have argued that this behavior comes from the bees’ natural inclination to abandon their hives at the first sign of a wildfire, Cornell University bee biologist Tom Seeley believes it may be exactly the opposite—honeybees may engorge themselves with honey so that they can survive inside the hive for a long period of time after a wildfire, which could easily burn all of the flowering plants that they depend on for nectar.
Whatever the reason, beekeepers have always taken advantage of the fact that honeybees become docile at the smell of smoke.
This article comes directly from content in the video series Why Insects Matter: Earth’s Most Essential Species. Watch it now, on Wondrium.
Bees Have Expandable Sacks

The anatomical feature that allows honeybees to fill up on honey is the crop. The crop is the frontmost section of an insect’s gut that, in honeybees and some other insects, functions like an expandable sack. The crop is located in the abdomen and is surrounded by muscles that relax to allow it to fill with honey but can also contract if the bee wants to share honey with another member of its hive.
Bees also use their crop to fill up with nectar when visiting flowers, which they later regurgitate inside the hive and fan with their wings to reduce the water content through evaporation, turning it into honey.
So the crop is fundamental to the lifestyle of the honeybee and, by extension, to the role that honeybees have played throughout human history.
How Bee Eyes Work
Bees are very good at locating flowers thanks to their acute visual abilities. Let’s take a closer look at one of the individual ommatidia that make up the compound eye of a honeybee.
An individual ommatidium is an elongated structure that tapers toward the base. At the top is the lens, which is made of two separate components. The outermost portion is the cornea, a clear, hexagonal structure. Below that is a crystalline cone, shaped like an upside-down pyramid.
Together, the cornea and cone focus light into the inner portion of the ommatidium called the rhabdom, which is lined by photoreceptor cells. Each of the photoreceptor cells inside an ommatidium is capable of responding at a particular wavelength of light.
A bee’s vision is also trichromatic like human vision, but the three colors that bees can see are different. Honeybees have photoreceptor cells attuned to green and blue as we do, but instead of red (which is a lower wavelength), bees have a third kind of photoreceptor cell that detects ultraviolet.
So compared to human vision, which can detect light of wavelengths between about 380 and 740 nanometers, honeybees can see wavelengths between 300 and 650 nanometers. So compared to us, a honeybee’s vision is effectively shifted towards shorter wavelengths of light.
Advantages of Bee Vision

Because bees are important pollinators for many plant species, the flowers of many bee-pollinated plants evolved to have colors that bees can easily see. This is the reason why so many bee-pollinated flowers are purple, violet, or blue. Some plants also have flowers with ultraviolet colors, which bees can see but are invisible to our eyes.
Using a photographic technique called ultraviolet-induced visible fluorescence photography, we can see flowers the way bees do. The petals and sepals appear to glow, but the parts of the flower most relevant to bees—the parts containing nectar and pollen—appear especially bright.
When the ultraviolet wavelengths are visible, some flowers have patterns that aren’t otherwise apparent. These are known as nectar guides because they help bees locate the precise place where the sugary reward for their services as pollinators can be found.
Bees Have Amazing Work Ethic
Their visual abilities make honeybees very efficient pollinators. Scientists estimate the 75% of the world’s flowering plants are pollinated by insects. They’re especially important for the plants we grow for food. Globally, about 80% of all crop plants are pollinated by insects, and of those, 92% are pollinated by bees.
In the United States, honeybees contribute $15 to $20 billion to the economy through their role as pollinators of crop plants. Some crops, like almonds, blueberries, and cherries, are almost totally dependent on honeybee pollination.
But the honeybee, Apis mellifera, isn’t native to North America. Honeybees were introduced from northern Europe in the early 1600s. Today beekeeping is a $5 billion industry, thanks to the role that honeybees play not only as crop pollinators but also in their production of honey and other products, like beeswax.
Common Questions about Why Beekeeping Is an Efficient Industry
One common theory behind this phenomenon is that this behavior comes from the bees’ natural inclination to abandon their hives at the first sign of a wildfire. The other theory is that the bees may engorge themselves with honey so that they can survive inside the hive for a long period of time after a wildfire, which could easily burn all of the flowering plants that they depend on for nectar.
The anatomical feature that allows bees to fill up on honey is the crop. It is located in the abdomen and functions like an expandable sack. Bees can fill it with honey, or with nectar when visiting flowers.
Beekeeping has turned into a profitable industry because of the important role bees play as pollinators. The production of honey and beeswax among other products has also helped the economy significantly.